Lycaenidae of Spain. Licénidos de España. Lepidoptera (Felipe Gil-T)

 Systematic List of Butterflies of Spain (Iberian Peninsula + Balearic + Canary islands) - List species - Lycaenidae of Spain

 

Total species

Nymphalidae

Lycaenidae

Pieridae

Papilionidae

Hesperiidae

Total

SPAIN (All/Todo)

100 79 28 5 31

243

 [* = species; ssp.= subspecies; var.= variety; syn.= synonym]

   [Papilionoidea - Hesperioidea]                          [taxon = in Iberia; taxon = Iberia & Balearics; taxon = Iberia & Canaries; taxon = Canaries; taxon = Iberia, Balearics & Canaries]

PAPILIONOIDEA Latreille, 1802    [Nymphalidae - Lycaenidae - Pieridae - Papilionidae]  

LYCAENIDAE [Leach, 1815]

Lycaenidae de España                                     Lycaenidae of Spain

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

Riodininae Grote, 1895  (1 species)    [Note: currently it is regarded as a different Family: Riodinidae]

Hamearis Hübner, [1819]:

* Hamearis lucina (L., 1758)

Theclinae Butler, 1869  (12 species)

Thecla Fabricius, 1807:

* Thecla betulae (L., 1758)

Favonius Sibatini & Ito, 1942 (= Neozephirus = Quercusia):

* Favonius quercus (L.., 1758): [ssp. iberica (Staundinger, 1901)

Tomares Rambur, 1840:

* Tomares ballus (Fabricius, 1787)

Laeosopis Rambur, 1858:

* Laeosopis roboris (Esper, [1793])

Satyrium Scudder, 1876:

* Satyrium w-album (Knoch, 1782)

* Satyrium spini ([Denis & Schiffermüller], 1775)

* Satyrium pruni (L., 1758)

* Satyrium acaciae (Fabricius, 1787)

* Satyrium ilicis (Esper, 1779)

* Satyrium esculi (Hübner, [1804]

Callophrys Billberg, 1820:

* Callophrys rubi (L., 1758)  [ssp. fervida  Staudinger, 1901]

* Callophrys avis Chapman, 1909

Lycaeninae [Leach, 1815]  (7 species)

Lycaena Fabricius, 1807:

* Lycaena tityrus (Poda, 1761)

* Lycaena (tityrus) bleusei (Oberthür, 1884)  [Note: this taxon is considered as species or subspecies according to authors]

* Lycaena virgaureae (L., 1758) [ssp. miegi (Vogel, 1857)]

* Lycaena phlaeas (L., 1761)

* Lycaena helle ([Denis & Schiffermüller], 1775)

* Lycaena hippothoe (L. 1761)

* Lycaena alciphron (Rottemburg, 1775) [ssp. gordius (Sulzer, 1776) (S. Spain: var. granatensis Ribbe, 1905)]

 

Polyommatinae Swainson, 1827  (59 especies)

            Lampides Hübner, [1819]:

                         * Lampides boeticus (L., 1767)

            Cacyreus Butler, 1897:

                         * Cacyreus marshalli Butler, 1898

            Leptotes Scudder, 1876:

                         * Leptotes pirithous (L., 1767)

            Tarucus Moore, 1881:

                         * Tarucus theophrastus (Fabricius, 1793)

            Zizeeria Chapman, 1910:

                         * Zizeeria knysna (Trimen, 1862)

            Azanus Moore, 1881:

                         * Azanus jesous (Guérin-Méneville, 1849) (in Algeciras, S. Cádiz prov. Probably introduced with ornamental Acacia spp.)

                         * Azanus ubaldus (Cramer, 1782) (in Gran Canaria and Fuerteventura islands. Probably introduced with ornamental Acacia spp.)

            Cupido Schrank, 1801:

                         * Cupido (Everes) argiades (Pallas, 1771)

                         * Cupido (Everes) alcetas (Hoffmannsegg, 1804)

                         * Cupido osiris (Meigen, 1829)

                         * Cupido carswelli Stempffer, 1927  (PDF) (PDF) (PDF) (PDF) : 

                               Note: recent molecular studies (DNA), 2015, have finally demonstrated that it is a different species to C. minimus.  (PDF) (PDF)

                                         (As Felipe Gil-T. said and affirmed in all his previously  published articles).

                         * Cupido lorquinii (Herrich-Schäffer, 1847)  (PDF) (PDF) (PDF)

                         * Cupido minimus (Fuessly, 1775)  (PDF) (PDF

            Cyclyrius Butler, 1897:

                         * Cyclyrius webbianus (Brullé, 1840)

            Celastrina Tutt, 1906:

                         * Celastrina argiolus (L., 1758)

            Pseudophilotes Beuret, 1958: 

                         * Pseudophilotes baton (Bergsträsser, 1779)

                         * Pseudophilotes panoptes (Hübner, 1813)

                         * Pseudophilotes abencerragus (Pierret, 1837)

            Scolitantides Hübner, [1819]:

                         * Scolitantides orion (Pallas, 1771)

            Glaucopsyche Scudder, 1872:

                         * Glaucopsyche alexis (Poda, 1761

                         * Glaucopsyche melanops (Boisduval, [1828]) [ssp. algirica Heyne, 1895:]

            Iolana Bethune-Baker, 1914:

                         * Iolana iolas (ssp?) (Ochsenheimer, 1816: [its final taxonomic status is pending of a future molecular study]: (PDF) (PDF)

            Maculinea Van Eecke, 1915:       

                                   [Note: some current authors consider to Maculinea as a synonym of the genus Phengaris Doherty, 1891, but other authors disagree]

                         * Maculinea arion (L., 1758)

                         * Maculinea nausithous (Bergsträsser, 1779)

                         * Maculinea alcon ([Denis & Schiffermüller], 1775) [ecological forms: alcon; rebeli (Hirschke, 1904)]

            Plebejus Kluk, 1780:

                         * Plebejus argus (L., 1758) [ssp. hypochionus (Rambur, 1858)]

                         * Plebejus idas (L., 1761) [ssp. nevadensis (Oberthür, 1896)]

            Kretania Beuret, 1959:               [Note: according to a recent molecular study, the "Plebejus pylaon" group is included in the genus Kretania]

                         * Kretania pylaon (= Plebejus pylaon) (Fischer von Waldheim, 1832):  

                                               The final taxonomic status of several taxa of the "pylaon" group is pending of molecular studies (DNA), like the next:

                                         Kretania p. hesperica (Rambur, 1839) (= Plebejus p. hespericus) (PDF) (PDF) 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

            Polyommatus Latreille, 1804:

                        * Polyommatus icarus (Rottemburg, 1775) [syn.= P. abdon Aistleiner & Aistleiner, 1994, according to molecular studies, DNA]

                        * Polyommatus celina (Austaut, 1879) [syn.= P. icarus celina] (DNA very different to P. icarus, genetic divergence > 5%)

                        * Polyommatus eros (Ochsenheimer, 1808)

                        * Polyommatus thersites (Cantener, 1835)

                        * Polyommatus amandus (Schneider, 1792) [spp. tova Higgins, 1948]

                        * Polyommatus escheri (Hübner, [1823])

                        * Polyommatus (Plebicula) golgus Plebicula) golgus (Hübner, 1813): (PDF) (PDF) (PDF) (PDF)

                        * Polyommatus (Plebicula) sagratrox (Aistleitner, 1986): Ecology-Morphology-Ethology-Distribution very different of P. golgus !!: (PDF) (PDF) (PDF) (PDF)

                        * Polyommatus (Plebicula) dorylas ([Denis & Schiffermüller], 1775)

                        * Polyommatus (Plebicula) nivescens (Keferstein, 1851)

                        * Polyommatus (Meleageria) daphnis ([Denis & Schiffermüller], 1775)

            Lysandra Hemming, 1933               (Note: Lysandra is a different genus to Polyommatus according to recent molecular studies, DNA)

                        * Lysandra coridon (Poda, 1761) [ssp. asturiensis Sagarra, 1924; f. syngraphoides]

                        * Lysandra caelestissima (Verity, 1921)

                        * Lysandra bellargus (Rottemburg, 1775) (PDF)

                        * Lysandra hispana (Herrich-Schäffer, 1852)

                        * Lysandra albicans (Gerhard, 1851): [ssp. albicans = a cline, gradual change; var. arragonensis Gerhard; var. bolivari Romei] (PDF)

            Agrodiaetus Hübner, 1822:          (Note: Agrodiaetus is regarded as a subgenus of Polyommatus by some recent authors)

                        * Agrodiaetus damon ([Denis & Schiffermüller], 1775) [ssp. noguerae (Sagarra, 1924)]

                        * Agrodiaetus ripartii (Freyer, 1830) (syn. "A.  agenjoi " Forster 1965)

                        * Agrodiaetus fabressei (Oberthür, 1910)

                        * Agrodiaetus fulgens Sagarra, 1925 [= a cline, only varities or forms. Synonyms: A. f. ainsae Foster, 1961 = A. ainsae pseudovirgilia = leonensis...]

                        * Agrodiaetus violetae Gómez-Bustillo, Expósito & Martínez, 1979 (PDF) (PDF) (PDF) (PDF) (PDF):

                              [ssp. violetae; ssp. subbaeticus Gil-T. & Gil-Uceda, 2005 (syn. A. fabressei subbaeticus). Also, the two sspp. are genetically different]

            Cyaniris Dalman, 1816:               (Note: Cyaniris is a different genus to Polyommatus according to several molecular studies on their DNA)

                         * Cyaniris semiargus (Rottemburg, 1775) [ssp. tartessus Gil-T. & Huertas, 2007: genetically different to all the taxa of "Cyaniris"!: (PDF)]

            Agriades Hübner, [1819]:

                         * Agriades zullichi (Hemming, 1933) (PDF)

                         * Agriades glandon (Prunner, 1798)

                         * Agriades pyrenaicus (Boisduval, 1840): [ssp. pyrenaicus; ssp. asturiensis (Oberthür, 1910)]

            Aricia [Reichenbach], 1817:

                         * Aricia agestis ([Denis & Schiffermüller], 1775) [only in NE. Iberian Peninsula, near the Pyrenees, N. Catalonia]

                         * Aricia cramera Eschscholtz, 1821 [very different genetically of A. agestis]

                         * Aricia artaxerxes (Fabricius, 1793): [ssp. montensis Verity, 1928: a different species or a subspecies of artaxerxes according to authors.

                                                                                    Note: the genetic distance (COI, barcode) between A. artaxerxes, sensu stricto, and the taxon "montensis" is of 1%]

                         * Aricia (Pseudoaricia) nicias (Meigen, [1829]

                         * Aricia morronensis (Ribbe, 1910)  [several subspecies described are only synonyms or varities] (PDF) (PDF) (PDF):

                               [ssp. morronensis; ssp. ramburi Verity, 1929; ssp. hesselbarthi Manley & Allcard, 1970; ssp. chapmani Ribbe, 1910]

            Eumedonia Forster, 1938:            (Note: Eumedonia  is a different genus to Aricia according to recent molecular studies on their DNA)

                         * Eumedonia eumedon (Esper, [1780]) (PDF) (PDF) (PDF)

                               [ssp. rondensis Gil-T. & Moreno-Benítez, 2012; ssp. axarquia (Gil-T., 2004); ssp. mariensis Eitschberger & Steiniger,1975]